1965 B.A. (Moderatorship in Genetics), Trinity College, Dublin University.
1970 Ph.D. Dublin University (TCD). (1967-1969 in University of Cincinnati) 1981 Sc.D. Dublin Univ. (External examiner, F. Sanger).
1969-71 EMBO Long Term Fellow, Dept. of Molecular Biology, University of Århus, Denmark
1971-72 I.C.R.F. Fellow, Cancer Research UK (present name), London.
1972-75 & ’77-‘78 Cold Spring Harbor Laboratory, Cold Spring Harbor, N.Y. 1975-77 Senior Scientific Officer ICI Corporate Lab (prior to the formation of Zeneca and AstraZeneca) Runcorn, England, on secondment to the Department of Molecular Biology, University of Edinburgh, for a Joint research scheme.
1978-80 Research Associate, Howard Hughes Medical Institute, U. of Utah.
1981 EMBO Short Term Fellow, Dept. of Molecular Biology, University of Geneva 1981-86 College and later Statutory Lecturer, Dept. of Biochemistry, UCC, Ireland.
1983 First Irish elected member of EMBO
1986- Research Associate Prof, then Research Prof. Department of Human Genetics, University of Utah – currently Emeritus status.
1997- Honorary Professor of Genetics, Department of Genetics, Trinity College Dublin, Ireland.
2002-03 1st Director of Life Sciences, Science Foundation Ireland.
2004- Research Professor, Biochemistry and also Microbiology, University College Cork, Ireland (later via UCC Academy DAC).
2007 First Royal Irish Academy gold medal for the Life Sciencaes.
2015 Member of review panel in Berlin of a Deutsche Forschungsgemeinschaft (DFG) Research Programme (1805/2).